Shadow Angels have been waiting for 12,000 years to put a death-blow to humanity, but as soon as life presented them such chance, an international organization Deava stands on their way. Appolo is a teenage pilot of one of Deava’s fighters that is the part of the robot Aquarion, an incarnation of a fallen angel Appolo. The 15-years-old boy’s fate is to oppose his eternal rival Touma in the decisive battle that will determine the fate of the Earth.