Ty, a one-of-a-kind designer running sneaker, lives a carefree life wrapped in absolute comfort inside his velvet box. Everything changes when his sister Maxine is snatched by a vain, ruthless Collector. In a frantic bid to save her, Ty gets lost on the loud, unpredictable streets of New York City. Stranded in the sprawling metropolis, the sheltered hero crosses paths with JB, a scuffed, street-smart sneaker. To rescue their loved ones, this unlikely pair of footwear friends must band together and step way out of their comfort zone.